When security fears forced Jordan to close its last border crossing with Syria this month, it severed its most important trade route to the outside, further straining an economy already hit by fallout from war.

But the damage to heavily import-reliant Jordan is not just hypothetical -- hundreds of importers and exporters lost millions of dollars worth of goods when rebels pillaged the vast free trade zone that straddled the border, most of it on the Syrian side.

Two Swedish men held hostage by Syrian jihadists have been reunited with their families in Sweden, the country's foreign ministry said on Sunday.

The hostages, named in the Swedish media as Thomas Olsson, 50, and Martin Reen, 33, were freed with the help of Palestinian and Jordanian authorities, said the ministry.

At least 92 people have been killed in Saudi-led coalition air strikes and fighting between supporters of President Abedrabbo Mansour Hadi and his opponents in southern Yemen, sources said Saturday.

In the key port city of Aden and in the adjacent Lahj province alone, 46 rebels died in ongoing clashes and 12 hours of intense air strikes that lasted until early Saturday, a military source close to them said.

Egyptian police said Saturday that four Islamists were killed in an accidental explosion while preparing bombs south of the capital.

Police said the suspected militants were members of the blacklisted Muslim Brotherhood movement of former Islamist president Mohamed Morsi, whom the army toppled in 2013.

The Islamic State group claimed an attack Saturday by French, Belgian and Senegalese suicide bombers on the Iraq-Jordan border that officials said killed at least four Iraqi security personnel.

The bombers detonated explosives-rigged vehicles, one of them a lorry, at checkpoints on the Iraqi side of the Turaibil crossing, said Sabah Karhout, the head of the Anbar province council.

The Islamic State jihadist group killed two senior officers and 10 other soldiers in a suicide bombing and heavy fighting in Iraq's Anbar province, a general said on Saturday.

"The commander of the 1st Division, Staff Brigadier General Hassan Abbas, and the commander of the 1st Brigade of the 1st Division, Staff Colonel Hilal Matar, and 10 soldiers were killed and 10 wounded," Staff Major General Mohammed al-Dulaimi said.

Fierce fighting raged Saturday in south Yemen between Iran-backed rebels and loyalists of exiled President Abedrabbo Mansour Hadi, as pressure mounted for the warring factions to hold political talks.

The Huthi rebels, who have overrun large parts of the country and forced Hadi to flee overseas, have demanded a complete end to a month of Saudi-led air strikes against them as a condition for U.N.-sponsored talks.

Al-Qaida's Syrian affiliate and its allies seized the last major government-held city in Idlib province on Saturday, in a blow that could expose the regime's coastal heartland to rebel attack.

The capture of Jisr al-Shughur in the northwestern province comes nearly a month after the same coalition of opposition forces, known as the "Army of Conquest", overran the provincial capital.

Israeli police shot dead a knife-wielding Palestinian who attempted to stab colleagues at an east Jerusalem checkpoint, a spokeswoman said on Saturday.

The 17-year-old assailant from the Al-Tur neighborhood of annexed Arab east Jerusalem managed to get past one checkpoint but was brought down at a second near Al-Zaim without any police casualties, the spokeswoman said.

The United Nations envoy to war-torn Syria said Friday that he will know by the end of June if there is any appetite among the opposing sides to find a peaceful way out of the conflict.

During closed discussions with the 15-member U.N. Security Council, Staffan de Mistura acknowledged that the "odds and conditions for launching a political transition are no better now than six months ago," diplomats at the briefing said.