Al-Qaida militants have killed at least 20 Shiite rebels in fresh clashes for control of parts of central Yemen, tribal sources said on Monday.
The rebels, known as Huthis, have been facing fierce resistance from al-Qaida fighters and Sunni tribesmen as they seek to expand their territory after seizing the capital Sanaa and the Red Sea port city of Hudeida.
Heaving fighting erupted overnight Sunday to Monday in the central town of Rada, a mixed Sunni-Shiite area that has been the scene of frequent clashes.
The rebels were killed in a car bombing that targeted a building where they had gathered and in subsequent clashes, tribal and security sources told Agence France Presse, adding that 12 rebels were also captured by al-Qaida militants.
The town was rocked by heavy explosions, with rocket-propelled grenades and artillery used by both sides in several hours of clashes, security officials said.
The al-Qaida militants also attacked rebel positions northeast of Rada and along a road connecting the town in Baida province to neighboring Dhamar, a Shiite-populated province taken last week by the rebels.
The Huthis have seized on chronic instability in Yemen since the 2012 ouster of veteran strongman Ali Abdullah Saleh to take control of large parts of the country.
President Abdrabuh Mansur Hadi's weak Sunni-led central government has failed to stop the rebels, despite a U.N.-brokered peace deal that was supposed to see them withdraw from the capital.
Al-Qaida in the Arabian Peninsula (AQAP), the extremist network's powerful Yemeni branch, and Sunni tribesmen have instead fought to halt the rebel advance, leaving dozens dead including 47 Huthi supporters in a suicide bombing in Sanaa this month.
- 'Failed' withdrawal talks -
The fighting has raised fears of Yemen -- located next to oil kingpin Saudi Arabia and important shipping routes in the Gulf of Aden -- collapsing into a failed state.
Hadi's government is also a key U.S. ally in the fight against al-Qaida, allowing Washington to conduct a longstanding drone war against the group on Yemeni territory.
The rebels faced no resistance when they took control of Sanaa last month and have refused to leave despite appearing to agree to the naming of a new prime minister under the U.N. deal.
They have since moved south and easily captured Dhamar but have faced heavy fighting in Sunni-majority Ibb province and from al-Qaida in Baida province, where Rada is located.
On Sunday, negotiations hosted by provincial governor Yehya al-Iryani to convince all fighters to withdraw from Ibb city "failed", one of the mediators attending the talks told AFP.
"The Huthis insisted on deploying their fighters alongside security forces in Ibb... to fight al-Qaida and prevent it from taking over the province," the source said.
The Shiite rebels have traditionally been concentrated in the northern parts of Yemen, on the border with Sunni-dominated Saudi Arabia.
Yemeni authorities and Saudi Arabia have accused Iran of backing the Huthis in a similar fashion to its support for Lebanon's powerful Shiite militia Hezbollah.
Gulf Arab states have warned that instability in Yemen is threatening regional security.
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